1. The University of Manchester, Manchester, United Kingdom; Achúcarro Basque Center for Neuroscience, IKERBASQUE, Basque Foundation for Science, Bilbao, Spain; Department of Neuroscience, University of the Basque Country UPV/EHU and CIBERNED, Leioa, Spain; Center for Basic and Translational Neuroscience, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark; and Center for Translational Neuromedicine, University of Rochester Medical Center, Rochester, New York