1. Forest and Rangeland Department, Khorasan Razavi Agricultural and Natural Resources Research and Education Center. AREEO . Mashhad , Iran .
2. Faculty of Biologia , Shahid Beheshti University , Tehran , Iran .
3. Research institute of Forests and Rangelands. AREEO , Tehran , Iran .
4. Department of biology , Payamenoor University , P. O. Box 19395-3697, Tehran , Iran .
5. Department of Biology School of Science, Tehran, Science and Research Branch , Islamic Azad University , Tehran , Iran