1. University of Debrecen , Department of Physical Geography and Geoinformatics , Egyetem tér 1, 4032 Debrecen , Hungary
2. MTA-DE Biodiversity and Ecosystem Services Research Group , Egyetem tér 1, 4032 Debrecen , Hungary
3. University of Debrecen , Department of Evolutionary Zoology and Human Biology , Egyetem tér 1, 4032 Debrecen , Hungary
4. MTA-DE Lendület Seed Ecology Research Group , Egyetem tér 1, 4032 Debrecen , Hungary
5. University of Debrecen , Department of Landscape Protection and Environmental Geography , Egyetem tér 1, 4032 Debrecen , Hungary