1. CSIC, Global Ecology Unit CREAF-CEAB-UAB, Cerdanyola del Vallès, 08193 Catalonia , Spain
2. Department of Agroforestry Science , Universidad de Sevilla , , Sevilla , Spain
3. Department of Plant Sciencies and Microbiology, Escuela Politécnica Superior de Orihuela , , Alicante , Spain
4. Dipartimento di Scienze Agrarie, Alimentari e Agroambientali . University of Pisa , Pisa , Italy
5. Plant & Food Research , Batchelar Road , Palmerston North , New Zealand