1. Behavioural Ecology Group, Department of Systematic Zoology and Ecology , Eötvös Loránd University , 1117 Budapest, Pázmány Péter sétány 1/C , Hungary
2. Department of Ethology , Eötvös Loránd University , 1117 Budapest, Pázmány Péter sétány 1/C , Hungary
3. Department of Anatomy, Cell- and Developmental Biology , Eötvös Loránd University , 1117 Budapest, Pázmány Péter sétány 1/C , Hungary