1. Department of Fisheries, Faculty of Fisheries and Environmental Sciences , Gorgan University of Agricultural Sciences and Natural Resources , Gorgan , Iran
2. Razi Vaccine and Serum Research Institute, Agricultural Research, Education and Extension Organization (AREEO) , Karaj , Iran
3. Department of Animal and Aquatic Sciences, Faculty of Agriculture , Chiang Mai University , Chiang Mai 50200 , Thailand
4. Science and Technology Research Institute , Chiang Mai University , 239 Huay Keaw Rd., Suthep, Muang, Chiang Mai 50200 , Thailand