1. Abdurakhmanov, Yu.A., Ryby presnykh vod Azerbaidzhana (Freshwater Fishes of Azerbaijan), Baku: Akad. Nauk AzSSR, 1962.
2. Barach, G.P., Fauna Gruzii, T. 1: Ryby presnykh vod (Fauna of Georgia, vol. 1: Freshwater Fishes), Tbilisi: Akad. Nauk GruzSSR, 1941.
3. Berg, L.S., Ryby presnykh vod SSSR i sopredel’nykh stran (Freshwater Fishes of the USSR and the Adjacent Countries), Moscow; Leningrad: Akad. Nauk SSSR, 1949, pt. 2, pp. 469–926.
4. Bobrinskii, N.A., Geografiya zhivotnykh. Kurs zoogeografii (Geography of Animals. Zoogeography Course), Moscow: Uchpedgiz, 1951.
5. Elanidze, R.F., Ikhtiofauna rek i ozer Gruzii (Ichthyofauna of Rivers and Lakes of Georgia), Tbilisi: Metsniereba, 1983.