1. Alekseyev, S.S., Sympatric forms of lenok (genus Brachymystax) in the Vitim River basin, Biol. Nauki, 1985, no. 3, pp. 41–48.
2. Alekseyev, S.S. and Kirillov, A.F., Morphology and distribution of two forms of lenok of genus Brachymystax Günther (Salmonidae) in the Lena River basin, Vopr. Ikhtiol., 1985, vol. 25, no. 4, pp. 597–602.
3. Alekseyev, S.S., Kirillov, A.F., and Samusenok, V.P., Distribution and morphology of the sharp-snouted and bluntsnouted lenoks of genus Brachymystax (Salmonidae) in Eastern Siberia, Vopr. Ikhtiol., 2003, vol. 43, no. 3, pp. 311–333.
4. Antonov, A.L. and Knizhin, I.B., Thymallidae fishes from Amur River basin: existing studies and modern diversity, Amur. Zool. Zh., 2014, vol. 6, no. 3, pp. 298–307.
5. Atlas presnovodnykh ryb Rossii (Atlas of Freshwater Fishes of Russia), Reshetnikov, Yu.S., Ed., Moscow: Nauka, 2003, vol. 1.