1. Andriyashev, A.P., Ryby severnykh morei SSSR (Fishes of the Northern Seas of the USSR), Moscow: Akad. Nauk SSSR, 1954.
2. Vasilyev, V.P., Evolyutsionnaya kariologiya ryb (Evolutionary Karyology of Fish), Moscow: Nauka, 1985.
3. Lindberg, G.U. and Krasyukova, Z.V., Ryby Yaponskogo morya i sopredel’nykh chastei Okhotskogo i Zheltogo morei (Fishes of the Sea of Japan and the Adjacent Parts of the Sea of Okhotsk and Yellow Sea), Leningrad: Nauka, 1987, part 5.
4. Moreva, I.N. and Borisenko, S.A., The karyotype of the flathead sculpin Megalocottus platycephalus platycephalus (Pallas, 1814) (Pisces: Cottidae) from Odyan Bay, Sea of Okhotsk, Russ. J. Mar. Biol., 2014, vol. 40, no. 2, pp. 125–130.
5. Moreva, I.N. and Borisenko, S.A., The karyotype of the flathead sculpin Megalocottus platycephalus taeniopterus (Kner, 1868) (Pisces: Cottidae) from Vostok Bay, Sea of Japan, Russ. J. Mar. Biol., 2015, vol. 41, no. 6, pp. 456–460.