1. College of Biology and the Environment, Nanjing Forestry University, Nanjing, Jiangsu, China,
2. Hangzhou Key Laboratory for Animal Adaptation and Evolution, School of Life Sciences, Hangzhou Normal University, Hangzhou, Zhejiang, China, and
3. Jiangsu Key Laboratory for Biodiversity and Biotechnology, College of Life Sciences, Nanjing Normal University, Nanjing, Jiangsu, China