1. Wern Veterinary Surgeons, Department of Research and Innovation, Unit 11, Lon Parcwr Industrial Estate, Ruthin, Denbighshire, LL15 1NJ, UK and Department of Epidemiology and Population Health, Institute of Infection and Global Health, University of Liverpool, Leahurst Campus, Neston, Wirral, Cheshire CH64 7TE
2. Institute of Veterinary Science, University of Liverpool, Leahurst Campus, Neston, Wirral, Cheshire CH64 7TE