1. Department of Clinical Veterinary Science, University of Bristol, Langford House, Langford, Bristol BS40 5DU, UK
2. Department of Pathology and Microbiology, School of Medical Sciences, University of Bristol, Bristol BS8 1TD, UK
3. Department of Pathology and Microbiology, University of Bristol, Langford House, Langford, Bristol BS40 5DU, UK