1. Laboratory of Virus Pathogenesis,
2. Center for Emerging Virus Research, and
3. Laboratory of Viral Control, Institute for Virus Research, Kyoto University, Kyoto, Japan;
4. Department of Tumor Virology, Institute for Genetic Medicine, Hokkaido University, Sapporo, Japan;
5. Pathology and Tumor Biology, Graduate School of Medicine, Kyoto University, Kyoto, Japan;
6. Doshisha Women's College of Liberal Arts, Kyotanabe, Japan;
7. Laboratory of Viral Oncology, Aichi Cancer Center, Nagoya, Japan; and
8. Central Institute for Experimental Animals, Kawasaki, Japan