1. Department of Immunology, Wright-Fleming Institute, Imperial College London, London, United Kingdom;
2. Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, PA;
3. Illumina, Chesterford Research Park, Essex, Little Chesterford, United Kingdom;
4. University of California, San Diego, CA; and
5. Department of Genitourinary Medicine and Communicable Diseases, Wright-Fleming Institute, Imperial College London, London, United Kingdom