1. Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA;
2. Department of Ophthalmology, Otorhinolaryngology, and Head and Neck Surgery, Ribeirao Preto School of Medicine, University of São Paulo, São Paulo, Brazil;
3. Medical Research Council Centre for Molecular Bacteriology and Infection, Imperial College London, London, United Kingdom;
4. Department of Pediatrics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA; and
5. Department of Microbiology, New York University School of Medicine, New York, NY