1. School of Immunity and Infection, Centre for Translational Inflammation Research, College of Medical and Dental Sciences, University of Birmingham, Birmingham, United Kingdom;
2. Transplantation Immunology Group, Department of Haematology, Cancer Institute and Institute of Immunity and Transplantation, University College London, London, United Kingdom;
3. Cancer Research UK Viral Oncology Group, University College London, London, United Kingdom; and
4. School of Clinical and Experimental Medicine, College of Medical and Dental Sciences, University of Birmingham, Birmingham, United Kingdom