1. Northern Ireland Centre for Food and Health, Centre for Molecular Biosciences, University of Ulster, Cromore Road, Coleraine, N. Ireland, UK
2. Environmental and Biochemical Sciences Group, Enhancing Crop Productivity and Utilisation Theme, The James Hutton Institute, Invergowrie, Dundee, Scotland, UK
3. School of Life Sciences, Heriot-Watt University, Edinburgh, Scotland, UK