1. Queen Square Brain Bank for Neurological Disorders, UCL Institute of Neurology, Queen Square, University College London, London, UK
2. Sobell Department of Motor Neuroscience and Movement Disorders, UCL Institute of Neurology, Queen Square, University College London, London, UK
3. Department of Neurology, Niigata University Brain Research Institute, Niigata, Japan
4. The Sara Koe PSP Research Centre, Department of Molecular Neuroscience, UCL Institute of Neurology, Queen Square, University College London, London, UK
5. Dementia Research Centre, UCL Institute of Neurology, Queen Square, University College London, London, UK
6. The Reta Lila Weston Institute of Neurological Sciences, Wakefield Street, UCL Institute of Neurology, Queen Square, University College London, London, UK
7. Department of Pathology, Niigata University Brain Research Institute, Niigata, Japan
8. Department of Molecular Neuroscience, Niigata University Brain Research Institute, Niigata, Japan
9. Department of Neuropathology, Institute of Brain Science, Hirosaki University School of Medicine, Hirosaki, Japan