1. Department of Neurology and Neurological Science, Graduate School of Medical and Dental Science, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan
2. Department of Neurology, Tokyo Metropolitan Institute for Neuroscience, Fuchu, Tokyo, Japan
3. Molecular Neuropathology, Tokyo Metropolitan Institute for Neuroscience, Fuchu, Tokyo, Japan
4. Centre for Research in Neurodegenerative Diseases, Departments of Medical Biophysics and Medicine (Neurology), University of Toronto, Toronto, Ontario, Canada
5. Department of Clinical Neuroscience, University of Cambridge, Cambridge, United Kingdom