1. Department of Zoology, Institute of Biosciences, University of São Paulo, 05508-090, São Paulo, SP, Brazil.
2. Amphibian Collection, Museum of Zoology of the University of São Paulo, 04263-000, São Paulo, SP, Brazil.
3. Universidad Autónoma de Chiriquí, David, Republic of Panama.
4. Los Naturalistas, David 0426-01459, Chiriquí, Panama.
5. Corporación Colombia en Hechos, Bogotá, Colombia.