1. CAS Key Laboratory of Coastal Environmental Processes and Ecological Remediation, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Key Laboratory of Coastal Environmental Processes, YICCAS, Yantai, Shandong 264003, P. R. China
2. University of Chinese Academy of Sciences, Beijing 100049, P. R. China
3. Laboratory for Marine Biology and Biotechnology, Qingdao Marine Science and Technology Center, Qingdao, Shandong 266237, P. R. China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, Shandong 266071, P. R. China