1. School of Earth and Ocean Sciences, Univeristy of Victoria, Victoria V8P 5C2, Canada
2. Institute of Ocean Sciences, Department of Fisheries and Oceans, 9860 West Saanich Road, Sidney, BC V8L 4B2, Canada
3. EAWAG – Swiss Federal Institute of Aquatic Science and Technology, Department Surface Waters, 6047 Kastanienbaum, Switzerland