1. Laboratory for Physical Biology, RIKEN Center for Biosystems Dynamics Research, Kobe 650-0047, Japan
2. Nonequilibrium Physics of Living Matter RIKEN Hakubi Research Team, RIKEN Center for Biosystems Dynamics Research, 2-2-3 Minatojima-minamimachi, Chuo-ku, Kobe 650-0047, Japan
3. Department of Physics, Emory University, Atlanta, GA, USA
4. Department of Physics, Syracuse University, Syracuse, New York 13244, USA
5. BioInspired Institute, Syracuse University, Syracuse, New York 13244, USA