1. Division of Food Sciences, School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, LE12 5RD, UK
2. School of Agriculture, Biomedicine and Environment, La Trobe University, Bundoora, Victoria, Australia, 3086
3. Division of Plant Sciences, School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, LE12 5RD, UK