1. College of Chemical and Biological Engineering, Shandong University of Science and Technology, Qingdao, 266590, P. R. China
2. Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, 266101, P. R. China
3. Department of Biology and Chemical Engineering, Qingdao Technical College, Qingdao, 266555, P. R. China