1. CAS Key Laboratory of Coastal Environmental Processes and Ecological Remediation, Shandong Key Laboratory of Coastal Environmental Processes, YICCAS, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS), Yantai 264003, Shandong, P. R. China
2. University of Chinese Academy of Sciences, Beijing 100049, P. R. China
3. Center for Ocean Mega-Science, Chinese Academy of Sciences Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, Shandong, P. R. China
4. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao 266237, Shandong, P. R. China