1. Fukushima Regional Collaborative Research Center, National Institute for Environmental Studies, 10-2 Fukasaku, Miharu, Fukushima 963-7700, Japan
2. Institute of Agriculture, Tokyo University of Agriculture and Technology, 3-5-8 Sauwau-cho, Fuchu, Tokyo 183-8509, Japan
3. Field Science Center for Northern Biosphere, Hokkaido University, 131 Toikanbetu, Horonobe-cho, Teshio-gun, Hokkaido 098-2943, Japan
4. Graduate School of Bioagricultural Sciences, Nagoya University, Furo-cho, Chikusa-ku, Nogoya, Aichi 464-8601, Japan