1. Key Laboratory of Coastal Biology and Bioresource Utilization, Yantai Institute of Coastal Zone Research, Chinese Academy of Sciences, Yantai 264003, Shandong, China
2. School of Resources and Environmental Engineering, Ludong University, Yantai 264025, Shandong, China
3. College of Life Sciences, Yantai University, Yantai 264003, Shandong, China