1. Key Laboratory of Horticultural Plant Biology, Ministry of Education/College of Horticulture and Forestry Sciences, Huazhong Agricultural University, Wuhan, China 430070
2. MOA Key Laboratory of Crop Ecophysiology and Farming System in the Middle Reaches of the Yangtze River, College of Plant Science & Technology, Huazhong Agricultural University, Wuhan, China 430070
3. Hubei Hongshan Laboratory, Wuhan, China 430070
4. College of Agronomy and Biotechnology, China Agricultural University, Beijing 100083, China