1. Institute of Microbial Engineering, Laboratory of Bioresource and Applied Microbiology, School of Life Sciences, Henan University, Kaifeng, 475004, People's Republic of China
2. Engineering Research Center for Applied Microbiology of Henan Province, Kaifeng 475004, People's Republic of China
3. School of Pharmacy, Henan University, Kaifeng, Henan, 475004, China