1. Department of Biology, Aarhus University Centre for Water Technology, Aarhus University, Section for Microbiology, Ny Munkegade 114, 8000 Aarhus C, Denmark
2. Department of Biological and Chemical Engineering, Aarhus University, Gustav Wieds Vej 10, 8000 Aarhus C, Denmark
3. Department of Agroecology, Pioneer Centre Land-CRAFT, Aarhus University, C. F. Møllers Allé 4, 8000 Aarhus C, Denmark
4. Department of Plant and Environmental Sciences, University of Copenhagen, Thorvaldsensvej 40, 1871 Frederiksberg, Denmark
5. School of Environmental Sciences, University of Guelph, Ontario N1G2W1, Canada