1. Key Laboratory of Life-Organic Analysis of Shandong Province
2. Qufu Normal University
3. Qufu 273165
4. China
5. Key Laboratory of Tibetan Medicine Research & Qinghai Key Laboratory of Qinghai-Tibet Plateau Biological Resources
6. Northwest Institute of Plateau Biology
7. Chinese Academy of Science
8. Xining 810001
9. School of Food and Biological Engineering