1. Institute of Natural Sciences, Shanghai Jiao Tong University, Shanghai 200240, China
2. Department of Cell and Developmental Biology & Michigan Neuroscience Institute, University of Michigan Medical School, 48105, Ann Arbor, MI, USA
3. Shanghai National Center for Applied Mathematics (SJTU Center), Shanghai Jiao Tong University, Shanghai 200240, China
4. State Key Laboratory for Microbial Metabolism, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai 200240, China
5. Zhangjiang Institute for Advanced Study, Shanghai Jiao Tong University, Shanghai 200240, China