1. Chongqing Key Laboratory of Industrial Fermentation Microorganism, Chongqing University of Science and Technology, Chongqing 401331, P. R. China
2. College of Chemistry and Chemical Engineering, Chongqing University of Science and Technology, Chongqing 401331, P. R. China
3. Fujian Key Laboratory on Conservation and Sustainable Utilization of Marine Biodiversity, Fuzhou Institute of Oceanography, Minjiang University, Fuzhou 350108, P. R. China