1. College of Ocean Food and Biological Engineering, Jimei University, No.43, Yindou Road, QiaoYing District, Xiamen, Fujian 361021, China
2. Research Center of Food Biotechnology of Xiamen City, Xiamen 361021, China
3. Fujian Provincial Key Laboratory of Food Microbiology and Enzyme Engineering, Xiamen 361021, China
4. Department of Food, Nutrition and Packaging Sciences, Clemson University, Clemson, SC 29634, USA