1. a Environmental and Life Sciences Graduate Program, Trent University, Peterborough ON K9J 7B8, Canada
2. d Department of Aquatic Sciences and Assessment, Swedish University of Agricultural Science, 750 07, Uppsala, Sweden
3. e *CWRA Member
4. b Environmental and Resource Studies, Trent University, Peterborough ON K9J 7B8, Canada
5. c Aquatic Research and Development Section, Ministry of Natural Resources, Peterborough ON K9J 7B8, Canada