1. Centre for Wildlife Ecology, Department of Biological Sciences, Simon Fraser University, Burnaby, British Columbia, Canada
2. Department of Forest and Conservation Sciences, University of British Columbia, Vancouver, British Columbia, Canada
3. Wildlife Research Division, Wildlife and Landscape Science Directorate, Environment and Climate Change Canada, Saskatoon, Saskatchewan, Canada
4. Department of Biology, Western University, London, Ontario, Canada
5. Wildlife Research Division, Wildlife and Landscape Science Directorate, Environment and Climate Change Canada, Delta, British Columbia, Canada