1. Department of Biology and Wildlife, University of Alaska Fairbanks, Fairbanks, Alaska, USA
2. Alaska SeaLife Center and School of Fisheries and Ocean Sciences, University of Alaska Fairbanks, Seward, Alaska, USA
3. U.S. Geological Survey, Alaska Science Center, Anchorage, Alaska, USA
4. U.S. Geological Survey, Alaska Cooperative Fish and Wildlife Research Unit, University of Alaska Fairbanks, Fairbanks, Alaska, USA