1. Program in Ecology, Evolution and Conservation Biology, Department of Natural Resources and Environmental Science, University of Nevada Reno, Reno, Nevada, USA
2. Department of Fish and Wildlife Conservation, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, USA
3. Department of Wildlife, Fisheries, and Conservation Biology, University of Maine, Orono, Maine, USA
4. Washington Department of Fish and Wildlife, Spokane Valley, Washington, USA