1. School of Marine and Environmental Sciences, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia
2. Eszterházy Károly University, Institute of Biology, Botany Department, Eger, Pf. 43, H-3301 Hungary
3. Muséum National d'Histoire Naturelle, Institut de Systématique, Évolution, Biodiversité, Case Postale 39, 57 rue Cuvier, 75231 Paris cedex 05, France
4. School of Environmental and Natural Resource Sciences, Universiti Kebangsaan Malaysia, 43600 Bangi, Selangor, Malaysia