1. Institute of Ecology, University of Georgia, Athens, Georgia, USA; †Australian Animal Health Laboratory, Commonwealth Scientific Industrial Research Organization, Geelong, Victoria, Australia; ‡School of Public Health and Tropical Medicine, James Cook University, Townsville, Queensland, Australia; §Institute of Zoology, Zoological Society of London, London, United Kingdom; and ¶National Wildlife Health Center, U.S. Geological Survey, Madison, Wisconsin, USA