1. Graduate School of Life and Environmental Sciences, University of Tsukuba; Tsukuba Japan
2. Center for Tsukuba Advanced Research Alliance, University of Tsukuba; Tsukuba Japan
3. PRESTO, JST; Saitama Japan
4. Anatomy and Embryology, Doctoral Program in Life System Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba; Tsukuba Japan