1. Developmental Biology Programme, European Molecular Biology Laboratory Meyerhofstrasse 1, PO Box 10.2209 69012 Heidelberg Germany
2. Department of Physiology, Programs in Genetics, Human Genetics, Neuroscience, and Developmental Biology, 513 Parnassus Avenue, University of California San Francisco California 94143 0444 USA
3. Department of Biochemistry, Programs in Genetics, Human Genetics, Neuroscience, and Developmental Biology, 513 Parnassus Avenue, University of California San Francisco California 94143 0444 USA