1. Institute for Molecular Bioscience, The University of Queensland Brisbane Queensland Australia
2. The Queensland Institute of Medical Research Brisbane Queensland Australia
3. Walter and Eliza Hall Institute of Medical Research Parkville Victoria Australia
4. School of Chemistry and Molecular Biosciences, Institute for Molecular Bioscience, The University of Queensland Brisbane Queensland Australia
5. Department of Internal Medicine, Carver College of Medicine, The University of Iowa Iowa City IA USA