1. Department of Ocean Science, The Hong Kong University of Science and Technology , Hong Kong, China
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) , Guangzhou 511458, China
3. Department of Bacteriology, University of Wisconsin–Madison , Madison, WI, USA
4. Department of Chemical and Biological Engineering, The Hong Kong University of Science and Technology , Hong Kong, China
5. Institute for Advanced Study, Shenzhen University , Shenzhen 518060, China