1. Department of Evolutionary Ecology, Institute of Organismic and Molecular Evolution, Johannes Gutenberg-University , Mainz, Germany
2. Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology , Tsukuba 305-8566, Japan
3. Department of Biological Sciences, Graduate School of Science, University of Tokyo , Tokyo 113-0033, Japan
4. Graduate School of Life and Environmental Sciences, University of Tsukuba , Tsukuba 305-8571, Japan
5. Department of Insect Symbiosis, Max-Planck-Institute for Chemical Ecology , Jena, Germany