1. State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing 100071, China
2. Utrecht Centre for Tick-borne Diseases (UCTD), Faculty of Veterinary MedicineUtrecht University, Yalelaan 1, Utrecht 3584 CL, The Netherlands
3. Department of Veterinary Tropical Diseases, Faculty of Veterinary ScienceUniversity of Pretoria, Private Bag X04, Onderstepoort 0110, South Africa