1. Centre for Infection and Inflammation Research, School of Medical Sciences, University of New South Wales Sydney New South Wales Australia
2. Department of Infectious Diseases, Prince of Wales Hospital Randwick New South Wales Australia
3. Viral Immunology Group, The Burnet Institute Melbourne Victoria Australia
4. Virology Division, SEALS, Prince of Wales Hospital Randwick New South Wales Australia
5. Department of Microbiology & Pathology, Beirne Carter Center for Immunology Research, University of Virginia Health Sciences Center Charlottesville VA USA