1. Graduate School of International Health Development, Nagasaki University, Nagasaki 852-8523, Japan
2. Institute of Tropical Medicine (NEKKEN), Nagasaki University, Nagasaki 852-8523, Japan
3. Independent Researcher in Liphuphwe, P.O. Box 180, Dedza, Malawi
4. Department of Biology, Chancellor College, The University of Malawi, P.O. Box 280, Zomba, Malawi