1. Institute of Infection and Immunity, Cardiff University School of Medicine Cardiff UK
2. Faculty of Health Sciences, University of Bristol Bristol UK
3. Mathematics Institute, University of Warwick Coventry UK
4. Institute of Cancer and Genetics, Cardiff University School of Medicine Cardiff UK
5. Cellular Immunology Laboratory, QIMR Berghofer Medical Research Institute Brisbane QLD Australia
6. Human Immunology Section, Vaccine Research Center, National Institute of Allergy and Infectious Diseases, National Institutes of Health Bethesda MD USA